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Adobe Acrobat XI Pro is a free program for creating, editing, and converting all kinds of PDF files. Its robust and effective tools ensure that you can create and convert thousands of documents quickly and easily.Histone Acetylation Promotes Developmental Cell-Specific Neurite Formation and Regulates Neuronal Survival.
Histone acetylation is a key process in chromatin remodeling during development, but it is unclear how acetylation of histone N-terminal tails affects neuronal differentiation. We found that depletion of HDAC4, an N-terminal lysine acetyltransferase, promotes neurite formation in P19 cells, in vivo, and in dissociated cortical neurons. Add-back of the deacetylase domain, but not the lysine acetyltransferase domain, was sufficient for the observed neurite-promoting activity. HDAC4 knockdown also suppressed apoptosis caused by irradiation in cultured cortical neurons. The transcription of BDNF, a marker for neuronal differentiation, was more induced in HDAC4-depleted neurons than in control neurons, whereas BDNF transcription levels in HDAC4-expressing neurons were diminished by treatment with inhibitors of acetyltransferases, including p300. These findings suggest that HDAC4-mediated histone acetylation influences the transcription of BDNF in developing cortical neurons. FILED
NOT FOR PUBLICATION NOV 13 2012

MOLLY C. DWYER, CLERK
d0c515b9f4

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